Forest ecology and management; Cork and cork oak modelling; Forest inventory; Forest biometry; New and traditional agroforestry systems; Statistical analysis; Rural development.


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Scientific Articles (ISI)

Marques, S., Rodrigues, A.R., Paulo, J.A., Botequim, B., Borges, J.G. (2024)

Addressing Carbon Storage in Forested Landscape Management Planning‚ÄĒAn Optimization Approach and Application in Northwest Portugal.

Forests 2024, 15, 408. . DOI:

Firmino, P. N., Tomé, M., Paulo, J. A. (2023)

Do Distance-Dependent Competition Indices Contribute to Improve Diameter and Total Height Tree Growth Prediction in Juvenile Cork Oak Plantations?

Forests 14(5), 1066. DOI:

Jorge C., Tomé M., Ruiz-Peinado R., Zribi L., Paulo J.A. (2023)

Quercus suber allometry in the West Mediterranean basin

Forests, 14(3): 649. . DOI:

Paulo J.A., Firmino P.N., Tomé M.A (2023)

A tree- and climate-dependent growth model to predict mature annual cork thickness under different climate change scenarios.

Modeling Earth Systems and Environment. . DOI:

Paulo, J. A., Santos, D. I. (2023)

Virgin cork colour and porosity as predictors for secondary cork industrial quality

Industrial Crops and Products 205 (1): 117513. . DOI:

Ferreiro-Domínguez N., Palma J.H.N., Paulo J.A., Rigueiro-Rodríguez A., Mosquera-Losada M.R (2022)

Assessment of soil carbon storage in three land use types of a semi-arid ecosystem in South Portugal.

Catena, 213: 106196. DOI:

Lecomte X., Paulo J.A., Tomé M., Veloso S., Firmino P.N., Faias S.P., Caldeira M.C. (2022)

Shrub understorey clearing and drought affects water status and growth of juvenile Quercus suber trees

Forest Ecology and Management, 503, 119760. DOI:

Paulo J.A., Firmino P.N., Faias S.P., Tomé M. (2021)

Quantile regression for modelling the impact of climate in cork growth quantiles in Portugal

European Journal of Forest Research, 140: 991‚Äď1004. DOI:

Faias S.P., Tomé M., Paulo J.A. (2020)

Inter-tree competition analysis in undebarked cork oak plantations as a support tool for management in Portugal

New Forests, 51(3): 489-505. DOI:

Paulo J.A., Villalobos R. (2020)

Quassia amara L. diameter growth under different shading conditions accessed by crown illumination index: implications for the management of Costa Rica natural and planted forests

Agroforestry Systems, 94(3): 761-778. DOI:

Rolo V., Hartel T., Aviron S., Berg S., Crous-Duran J., Franca A., Mirck J., Palma J.H.N., Pantera A. Paulo J.A., Pulido F.J., Seddaiu G., Thenail C., Varga A., Viaud V., Burgess P.J., Moreno G. (2020)

Challenges and innovations for improving the sustainability of European agroforestry systems of high nature and cultural value: stakeholder perspectives

Sustainability Science, 15(5): 1301-1315. DOI:

Aubard V., Paulo J.A., Silva J.M.N. (2019)

Long-term monitoring of cork and holm oak stands productivity in Portugal with Landsat imagery

Remote Sensing, 11(5): 525. DOI:

Crous-Duran J., Graves A., Paulo J.A., Mirck J., Oliveira T.S., Kaya S., García de Jalón S., Palma J.H.N. (2019)

Modelling tree density effects on provisioning ecosystem services in Europe

Agroforestry Systems, 93, 1985‚Äď2007. DOI:

Crous-Duran J., Graves A.R., Garcia-de-Jalón S., Paulo J.A., Tomé M., Palma J.H.N. (2019)

Assessing food sustainable intensification potential of agroforestry using a carbon balance method

iForest, 12: 85-91. DOI:

Faias S.P., Paulo J.A., Firmino P.N., Tomé M. (2019)

Drivers for annual cork growth under two understory management alternatives on a podzolic cork oak stand

Forests, 10(2): 133.

Faias, S. P., Paulo, J. A., Tomé, M. (2019)

Competition pattern in Portuguese young cork oak plantations

New Forests 51 (3): 489-505. DOI:

Gschwantner T., Alberdi I., Balazs A., Bauwens S., Bender S., Borota D., Bosela M., Bouriaud O., Canellas I., Donis J., Freudenschuss A., Herve J.C., Hladnik D., Jansons J., Kolozs L., Korhonen K.T., Kucera M., Kulbokas G., Kuliesis A., Lanz A., Lejeune P., Lind T., Marin G., Morneau F., Nagy D., Nord-Larsen T., Nunes L., Pantic D., Paulo J.A., Pikula T., Redmond J., Rego F.C., Riedel T., Saint-Andre L., Seben V., Sims A., Skudnik M., Solti G., Tomter S.M., Twomey M., Westerlund B., Zell J. (2019)

Harmonisation of stem volume estimates in European National Forest Inventories

Annals of Forest Science, 76(1): 24.

de Jalón S.G., Burgess P.J., Graves A., Moreno G., McAdam J., Pottier E., Novak S., Bondesan V., Mosquera-Losada R., Crous-Duran J., Palma J.H.N., Paulo J.A., Oliveira T.S., Cirou E., Hannachi Y., Pantera A., Wartelle R., Kay S., Malignier N., Lerberghe P.V., Tsonkova P., Mirck J., Rois M., Kongsted A.G., Thenail C., Luske B., Berg S., Gosme M., Vityi A. (2018)

How is agroforestry perceived in Europe? An assessment of positive and negative aspects by stakeholders

Agroforestry Systems, 92(4): 829-848. DOI:

Demertzi M., Paulo J.A., Faias S.P., Arroja L., Dias A.C. (2018)

Evaluating the carbon footprint of the cork sector with a dynamic approach including biogenic carbon flows

International Journal of Life Cycle Assessment, 23(7): 1448-1459. DOI:

Faias S.P., Paulo J.A., Palma J.H.N., Tomé M. (2018)

Understory effect on tree and cork growth in cork oak woodlands

Forest Systems, 27(1).

Moreno G., Aviron S., Berg S., Crous-Duran J., Franca A., de Jalón S.G., Hartel T., Mirck J., Pantera A., Palma J.H.N., Paulo J.A., Re G.A., Sanna F., Thenail C., Varga A., Viaud V., Burgess P.J. (2018)

Agroforestry systems of high nature and cultural value in Europe: provision of commercial goods and other ecosystem services

Agroforestry Systems, 92(4): 877‚Äď891.

Palma J.H.N., Crous-Duran J., Graves A., García de Jálon S., Upson M., Oliveira T., Paulo J.A., Ferreiro-Dominguez N., Moreno G., Burgess P. (2018)

Integrating belowground carbon dynamics into Yield-SAFE, a parameter sparse agroforestry model

Agroforestry Systems, 92 (4): 1047-1057.

Rois-D√≠az M., Lovrińá N., Lovrińá M., Ferreiro-Dom√≠nguez N., Mosquera-Losada M.R., den Herder M., Graves A., Palma J.H.N., Paulo J.A., Pisanelli A., Smith J., Moreno G., Garc√≠a S., Varga A., Pantera A., Mirck J., Burgess P. (2018)

Farmers’ reasoning behind the uptake of agroforestry practices: evidence from multiple case-studies across Europe

Agroforesty Systems, 92(4): 811-828.

Demertzi M., Sierra-Pérez J., Paulo J.A., Arroja L., Dias A.C. (2017)

Environmental performance of expanded cork slab and granules through life cycle assessment

Journal of Cleaner Production, 145: 294‚Äď302. DOI:

den Herder M., Moreno G., Mosquera-Losada R., Palma J.H.N, Sidiropoulou A., Freijanes J.J.S., Crous-Duran J., Paulo J.A., Tomé M., Pantera A., Papanastasis V.P., Mantzanas K., Pachana P., Papadopoulos A., Plieninger T., Burgess P.J. (2017)

Current extent and stratification of agroforestry in the European Union

Agriculture, Ecosystems and Environment, 241: 121‚Äď132. DOI:

Paulo J.A., Pereira H., Tomé M. (2017)

Analysis of variables influencing tree cork caliper in two consecutive cork extractions using cork growth index modelling

Agroforestry Systems, 91(2): 221‚Äď237.

Paulo J.A., Tomé M. (2017)

Using the SUBER model for assessing the impact of cork debarking rotation on equivalent annual annuity in Portuguese stands

Forest Systems, 26(1): e008.

Paulo J.A., Tomé M. (2017)

Does debarking intensity in the first cork extraction affect future cork thickness?

Annals of Forest Science, 74: 66. DOI:

Reyer C., Bathgate S., Blennow K., Borges J.G., Bugmann H., Delzon S., Faias S.P., Garcia-Gonzalo J., Gardiner B., Gonzalez-Olabarria J.R., Gracia C., Guerra J., Kellomäki S., Kramer K., Lexer M.J., Lindner M., van der Maaten E., Maroschek M., Muys B., Nicoll B., Palahi M., Palma J.H.N., Paulo J.A., Peltola H., Pukkala T., Rammer W., Ray D., Sabaté S., Schelhaas M., Seidl R., Temperli C., Tomé M., Yousefpour R., Zimmermann N.E., Hanewinkel M. (2017)

Are forest disturbances amplifying or canceling out climate change-induced productivity changes in European forests?

Environmental Research Letters, 12(3).

Demertzi M., Paulo J.A., Arroja L., Dias A.C. (2016)

A carbon footprint simulation model for the cork oak sector

Science of the Total Environment, 566‚Äď567:499‚Äď511. DOI:

Paulo J.A., Faias S.P., Ventura-Giroux C., Tomé M. (2016)

Estimation of stand crown cover using a generalized crown diameter model: application for the analysis of Portuguese cork oak stands stocking evolution

iForest – Biogeosciences and Forestry, 9: 437-444.

Palma J.H.N., Paulo J.A., Faias S.P., Garcia-Gonzalo J., Borges J.G., Tomé M. (2015)

Adaptive management and debarking schedule optimization of Quercus suber L. stands, under climate change. Case study in Chamusca, Portugal

Regional Environmental Change, 15:(8): 1569-1580.

Paulo J.A., Faias S., Gomes A.A., Palma J.H.N., Tomé J., Tomé M. (2015)

Predicting site index from climate and soil variables for cork oak (Quercus suber L.) stands in Portugal

New Forests, 46(2):293-307.

Paulo, J. A., Oliveira, V. Pereira, H. (2015)

Book of abstracts of the 2nd Centro de Estudos Florestais (CEF) conference from research to practice.

Cork oak stands and cork oak. 25th September 2015. Instituto Superior de Agronomia, Lisboa, Portugal. 27 pp. . URL:

Palma J.H.N., Paulo J.A., Tomé M. (2014)

Carbon sequestration of modern Quercus suber L. silvoarable agroforestry systems in Portugal: a YieldSAFE-based estimation

Agroforestry Systems, 88(5): 791-801.

Raftoyannis Y., Nocentini S., Marchi E., Calama Sainz R., Garcia Guemes C., Pilas I., Peric S., Paulo J.A., Moreira-Marcelino A.C., Costa-Ferreira M., Kakouris E., Lindner M. (2014)

Perceptions of forest experts on climate change and fire management in European Mediterranean forests

iForest, 7: 33-41. DOI:

Coelho M.B., Paulo J.A., Palma J.H.N., Tomé M. (2012)

Contribution of cork oak plantations installed after 1990 in Portugal to the Kyoto commitments and to the landowners economy

Forest Policy and Economics, 17: 59-68.

Faias S.P., Palma J.H.N., Barreiro S., Paulo J.A., Tomé M. (2012)

sIMfLOR ‚Äď Platform for the Portuguese forest simulators

Forest Systems, 21 (3): 543-548.

Paulo J.A., Tomé J., Tomé M. (2011)

Nonlinear fixed and random generalized height-diameter models for Portuguese cork oak stands

Annals of Forest Science, 68(2): 295-309.

Paulo J.A., Tomé M. (2010)

Predicting mature cork biomass with t years of growth from one measurement taken at any other age

Forest Ecology and Management, 259: 1993-2005.


Estudo prospetivo para o setor florestal

Caldeira B., Praxedes J., Santos P.M., Brígido S., Paulo J.A., Palma J.H.N., Pina J.P., Garcia-Gonzalo J., Borges J.G.C., Tomé M., Soares P., Barreiro S., Mendes A.M.S.C., Sottomayor M. (2013)

Associa√ß√£o para a Competitividade da Ind√ļstria da Fileira Floresta, 295 p.. URL:

Book of abstracts of the 1st Centro de Estudos Florestais (CEF) conference from research to practice.

Paulo, J. A., Pereira, H. (2013)

Cork oak stands and cork oak. 31st January 2014. Instituto Superior de Agronomia, Lisboa, Portugal. 17 pp. . URL:

Book Chapters

Tom√©, M., Ca√Īellas, I., Bonet, J. A., Paulo, J. A., Palma, J. H. N., Mart√≠nez de Arag√≥n, J., Miina, J., Sheppard, J., Kurttila, M., Sanch√©s-Gonzalez, M., Pasalodos, M., de-Miguel, Sergio. (2019)

Resource and management ‚Äď Novel management concepts to boost product diversity and secure higher product flows

In: Wolfslehner, B., Prokofieva, I. and Mavsar, R. (editors). Non-wood forest products in Europe: Seeing the forest around the trees. What Science Can Tell Us 10. European Forest Institute. pp 114. . URL:

Tomé, M., Sheppard, J., Alma, M. H., Carrasquinho, I., Christensen, C. J., Hellström, J., Kurttila, M., Marnila, P., Mattila, P., Miina, J., Mutke, S., Paulo, J. A., Palma, J. H., Pihlava, J., Pignatti, G., Sanchéz-González, M., Silva, C. S., Soares, P., Vanhanen, H. (2019)

Non-wood tree products in Europe.

In: Harald Vacik, Mike Hale, Heinrich Spiecker, Davide Pettenella & Margarida Tomé (Eds.) 2020: Non-Wood Forest Products in Europe, Ecology and management of mushrooms, tree products, understory plants and animal products, Outcomes of the COST Action FP1203 on European NWFPs, BoD, Norderstedt, 416p.. ISBN: 978-3-7494-7546-9

Ca√Īellas I., S√°nchez-Gonz√°lez M., Bogino S.M., Adame P., Moreno-Fern√°ndez D., Herrero C., Roig S., Tom√© M., Paulo J.A., Bravo F. (2017)

Carbon sequestration in Mediterranean oak forests

In: Managing Forest Ecosystems: the Challenge of Climate Change. Bravo F., LeMay V., Jandl R. (Eds.). Managing Forest Ecosystems, Vol. 34, Part V, pp. 403-427. Springer, Cham. ISBN: 978-3-319-28248-0



Landscape resilience knowledge alliance for agriculture and forestry in the Mediterranean basin

Duration: 2022-2024

CEF Budget: 329187.50‚ā¨ ‚ā¨


Impulso à sustentabilidade da floresta de carvalhos do Parque Natural de Montesinho através da inovação: valorização da bolota e da melada

Duration: 2021-2024

CEF Budget: 249.976,88 ‚ā¨


Tecnologias inovadoras & solu√ß√Ķes s√≥cio-ecol√≥gicas-econ√≥micas para territ√≥rios resistentes aos inc√™ndios na Europa

Duration: 2021-2025

CEF Budget: 633455.84 ‚ā¨ ‚ā¨


Gest√£o integrada e inteligente de florestas complexas e planta√ß√Ķes mistas no SUDOE

Duration: 2020-2023

CEF Budget: 149931,27‚ā¨ ‚ā¨


Eating the wild: Improving the value-chain of Mediterranean Wild Food Products

Duration: 2020-2023

CEF Budget: 67000 ‚ā¨


Agroforestry Innovation Networks

Duration: 2017-2019

CEF Budget: 256.701,80 ‚ā¨


Redes de Inova√ß√£o de Corti√ßa, Resinas e Comest√≠veis da bacia Mediterr√Ęnica

Duration: 2017-2020

CEF Budget: 158.063 ‚ā¨


Estudo e modelação da interação de sobreiros com a vegetação circundante

Duration: 2016-2019

CEF Budget: 156.000 ‚ā¨


Models and decision SUpport tools for integrated FOrest policy development under global change and associated Risk and UNcertainty

Duration: 2016-2020

CEF Budget: 275333.00 ‚ā¨


AGroFORestry that Will Advance Rural Development

Duration: 2014-2017

CEF Budget: 409.852 ‚ā¨